The first Mesoamerican civilizations began

right after people crossed the Bering land bridge
when nomadic hunters settled in the Valley of Mexico
once people learned how to farm in the tropical forest.
when the Maya built the first cities in Central America.

The first Mesoamerican civilizations began when the Maya built the first cities in Central America.

The first Mesoamerican civilizations began when the Maya built the first cities in Central America.

The first Mesoamerican civilizations began when nomadic hunters settled in the Valley of Mexico.

To arrive at this answer, we can go through each option and analyze its validity.

Firstly, the option "right after people crossed the Bering land bridge" is incorrect. The Bering land bridge was a landmass that once connected Asia and North America, allowing for the migration of early human populations. While this land bridge played a significant role in the peopling of the Americas, it is not directly related to the development of the first Mesoamerican civilizations.

Secondly, the option "once people learned how to farm in the tropical forest" is also incorrect. While agriculture played a crucial role in the development of Mesoamerican civilizations, it was not the primary factor that led to their formation. The knowledge of farming techniques came later in the development of these civilizations.

Lastly, the option "when the Maya built the first cities in Central America" is also incorrect. The Maya civilization, although one of the earliest and most influential Mesoamerican civilizations, did not build the first cities or mark the beginning of Mesoamerican civilizations as a whole.

Therefore, the accurate answer is "when nomadic hunters settled in the Valley of Mexico." This settlement marked the transition from a nomadic lifestyle to a sedentary one, leading to the formation of the first Mesoamerican civilizations in the region.